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In addition, BRs modulate many physiological activities such as photosynthetic functioning (i.e., chlorophyll content and photosynthetic capacity), antioxidant enzyme activities, and metabolism of carbohydrates and mitigate the negative effects of HMs on plants ( Other PGRs such as JA and SA (o-hydroxybenzoic acid) also relieve a range of environmental stresses, like HMs, including BRs (Wasternack, 2014